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C4 Plants

C4 plants have a mechanism to deal with photorespiration: they physically separate the light and dark reactions.
  • The light-dependent reactions occur in the mesophyll.
  • The Calvin cycle occurs in the bundle sheath cells.
  • The result is no build up of O2 near Rubisco.
Photo by Kelvinsong / CC BY

Reminder of Light-Dependent Reactions




The Strategy of C4 Plants

  • CO2 fixation occurs in the mesophyll by PEP carboxylase to produce oxaloacetate.
  • Oxaloacetate is converted to malate and transported to the bundle sheath cells.
  • Malate is broken down to release CO2 which then feeds into the Calvin cycle.
  • A 3-carbon molecule called pyruvate is returned to the mesophyll and converted back to PEP.


Examples of C4 Plants:

Usually found in hot, dry climates: sugar cane, certain types of grass, maize (corn).



Which of the following is true about the mechanism that C4 plants use to prevent photorespiration?